What led to the rediscovery of Mendel's laws of heredity and the development of the theory of mutation?

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His study of discontinuous variations especially in the evening primrose led to his rediscovery (reported in 1900) of Mendel's laws of heredity and to the development of the theory of mutation which he expounded in The Mutation Theory (1901-3 tr. 1909-10) and in Plant-Breeding (1907).

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