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The insect undergoes incomplete metamorphosis , the egg hatching directly into an aquatic naiad, or nymph, with chewing mouthparts, which passes through some 20 nymphal stages over a period of two years or more, feeding on algae and diatoms and breathing oxygen taken directly from water by gills. It emerges from the water to transform into a subadult phase known as the subimago, unique among insects, in which it has wings and can fly but has immature legs, tail, and reproductive system.
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